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Triplax festiva (Erotylidae), a pleasing fungus beetle

Most of the references cited below contain original descriptions of species having type localities within GSMNP or references to species occurring in the park that are rare or otherwise unusual. References to source data used in creating the GSMNP beetle checklist are also included. The list is far from exhaustive and suggestions for additional citations are welcome. 

Allen, R. T. 1980. A review of the subtribe Myadi: description of a new genus and species, phylogenetic relationships, and biogeography (Coleoptera: Carabidae: Pterostichini). Coleopterists Bulletin 34: 1-29.

Barr, T. C., Jr. 1962. The genus Trechus (Coleoptera: Carabidae: Trechini) in the southern Appalachians. Coleopterists Bulletin 19: 65-92.

Barr, T. C., Jr. 1969. Evolution of the Coleoptera (Carabidae) in the Southern Appalachians, pp. 67-92 in The Distributional History of the Biota of the Southern Appalachians Part I: Invertebrates (P. C. Holt, ed.). Research Division Monograph 1 Virginia Polytechnic Institute, Blacksburg, VA.

Barr, T. C., Jr. 1974. The eyeless beetles of the genus Arianops Brendel (Coleoptera, Pselaphidae). Bulletin of the American Museum of Natural History 154, 52 pp.

Barr, T. C., Jr. 1995. Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239-248.

Beal, R. S., Jr. 1985. A taxonomic revision of the Nearctic species of Cryptorhopalum (Dermestidae: Coleoptera). Transaction of the American Entomological Society 111: 171-221. 

Campbell, J. M. 1969. A revision of the New World Oxyporinae (Coleoptera: Staphylinidae). Canadian Entomologist 101: 225-268. 

Campbell, J. M. 1976. A revision of the genus Sepidophilus Gistel  (Coleoptera: Staphylinidae) of America north of Mexico. Memoirs of the Entomological Society of Canada 99: 1-89. 

Campbell, J. M. 1982. A revision of the genus Lordithon Thomson (Coleoptera: Staphylinidae) of North and Central America. Memoirs of the Entomological Society of Canada 119: 1-116. 

CMN, H. and A. Howden 2002. Records collected from the Howdens during a visit to the Canadian Museum of Nature by A. R. Cline, March 2002.

Carlton, C. E., and R. S. Anderson 2004. Occurrence of the Introduced Weevil Myosides seriehispidus Roelofs in Great Smoky Mountains National Park (Coleoptera: Curculionidae). Coleopterists Bulletin 58: 343

Carlton, C. E., and D. S. Chandler. Revision of the Nearctic genus Pseudactium (Coleoptera: Pselaphidae: Euplectinae). Coleopterists Bulletin 48: 171-190. 

Ciegler, J. C. 2000. Ground Beetles and Wrinkled Bark Beetles of South Carolina (Coleoptera: Geadephaga: Carabidae and Rhysodidae ). Biota of South Carolina, Volume 1. Clemson University. 149 pp.

Ciegler, J., and W. Merritt 2001. (Unpublished checklist provided to park by authors.)

Cornell, J. F., and  Puthz, V.  1997. Two new North American Stenus (Parastenus) species from the Appalachians (Insecta: Coleoptera, Staphylinidae: Steninae). Philippia 8: 131-136.

Dajoz, R. and A. 2002. (Unpublished checklist provided to park by authors.)

Ford, E. J. 1973.  A revision of the genus Petalium LeConte in the United States, Greater Antilles, and the Bahamas (Coleoptera: Anobiidae). USDA Technical Bulletin No. 1467, 40 pp.

Green, J. W.  1956. Revision of the Nearctic species of Photinus (Lampyridae: Coleoptera).  Proceedings of the California Academy of Sciences 28: 561-613.

Grigarick, A. A., and R. O. Schuster. 1971. A revision of Actium Casey and Actiastes Casey (Coleoptera: Pselaphidae).  University of California Publications in Entomology Vol. 67. 56 pp.

Gusarov, V. 2002. A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 81, 88 pp.

Gusarov, V. 2002. A revision of Nearctic species of the genus Earota Mulsant and Rey, 1874 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 92, 16 pp.

Gusarov, V. 2003. A revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synomic notes. Zootaxa 353, 134 pp.

Harpootlian, P. J. 2001. Scarab Beetles (Coleoptera: Scarabaeidae) of South Carolina. Biota of South Carolina, Volume 2. Clemson University. 157 pp.

Herman, L. H., Jr. 1975. Revision and phylogeny of the monogeneric subfamily Pseudopsinae for the World (Staphylinidae, Coleoptera). Bulletin of the American Museum of Natural History 155: 243-317.

Kavanaugh, D. H. 1979. Studies on the Nebriini (Coleoptera: Carabidae), III. New nearctic Nebria species and subspecies, nomenclatural notes, and lectotype designations. Proceedings of the California Academy of Sciences 42: 87-133.

Knull, J. N. 1949. Three new species of Cleridae (Coleoptera). Ohio Journal of Science 49: 199-200.

Konstantinov, A., and A. Tishechkin. 2004.  The First Nearctic Leaf Litter Flea Beetle (Coleoptera: Chrysomelidae) from the Great Smoky Mountains National Park. Coleopterists Bulletin 58: 71-76

Larson, D. J., Y. Alarie, and R. E. Roughley. 2000. Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, Ontario, Canada xiv + 982 pp.

Larson, D. J. and R. W. Wolfe. 1998. Revision of North American Agabus Leach (Coleoptera: Dytiscidae): the semivittatus-group. The Canadian Entomologist 130:27-54.

Lawrence, J. F., and A. F. Newton, Jr. 1975. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names), pp. 779-1006 in Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80^th Birthday of Roy A. Crowson (J. Pakaluk and S. A. Slipinski (eds.), Muzeum I Instytut Zoologii, Warsaw.

Lawrence, J. F., and K. Stephan. 1975. The North American Cerylonidae (Coleoptera: Clavicornia). Psyche 82: 131-166.

Löbl, I., and F. G. Calame. 1996. Taxonomy and Phylogeny of the Dasycerinae (Coleoptera: staphylinidae).  Journal of  Natural  History 30: 247-291.

Mayor, A. J. 2004.  Range Extension for Enoclerus muttkowskii (Wolcott) (Coleoptera: Cleridae) from Great Smoky Mountains National Park. Coleopterists Bulletin 58: 344.

Smetana, A. 1995. Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae). Classification, phylogeny and taxonomic revision. Memoirs on Entomology International 3, x+946 pp.

Park List 2001.  (Unpublished checklist derived from identified material in park collection.)

Park, O. 1953. New or little known species of pselaphid beetles of the United States, with observations on taxonomy and evolution of the family Pselaphidae. Bulletin of the Chicago Academy of Sciences 9: 249-283.

Park, O. 1956. New or little known species of pselaphid beetles from southeastern United States. Journal of the Tennessee Academy of Science 31: 55-100.

Park, O. 1965. Revision of the genus Batriasymmodes (Coleoptera: Pselaphidae). Transactions of the American Microscopical Society 84: 184-201.

Peck, S. B. 1973. A systematic revision and the evolutionary biology of the Ptomophagus (Adelops) beetles of North America (Coleoptera; Leiodidae; Catopinae), with emphsis on cave-inhabiting species. Bulletin of the Museum of Comparative Zoology 145, no. 2: 29-162..

Ravensford Project. 2002 checklist. Coleoptera collected from litter and soil samples. Identifications by Adreian Mayor. Specimens deposited in the University of Tennessee Museum. List provided by Dr. Ernie Bernard. 

Riley, E. G., and S. Clark.  Checklist of chrysomelid beetles collected and identified during the course of their research. Vouchers deposited in Texas A&M University Insect Collection. 

Selander, R. B. 1960. Bionomics, Systematics, and Phylogeny of Lytta, a Genus of Blister Beetles (Coleoptera: Meloidae). Illinois Biological Monographs: Number 28. 295 pp.

Sokolov, M., C. E. Carlton and J. F. Cornell. 2004. Review of Anillinus, with Descriptions of 17 New Species and a Key to Soil and Litter Species (Coleoptera: Carabidae: Trechinae: Bembidiinae). Coleopterists Bulletin 58: 185-233

Tremont. 2001. Tremont Institute, identifications sent by Paul Super.

Trieff, D. D. 2002.  Composition of the Coleoptera and associated insects collected by canopy fogging of Northern Red Oak (Quercus rubra L.) trees in the Great Smoky Mountains National Park and the University of Tennessee Arboretum.  Unpublished master's thesis, University of Tennessee.  86 pp.

Vit, S. 1995. Deux especes nouvelles d’Eucinetidae d’Amerique du Nord particulierement interessantes (Coleoptera: Eucinetidae). Elytron 9: 125-137.

Watrous, L. 1980. Lathrobium (Tetartopeus): natural history, phylogeny and revision of the nearctic species (Coleoptera, Staphylinidae). Systematic Entomology 5: 303-338.

Wheeler, Q. D., and J. V. McHugh. 1994. A new southern Appalachian species Dasycerus bicolor (Coleoptera: Staphylinidae: Dasycerinae) from declining endemic fir forests. Coleopterists Bulletin 48: 265-271.

Wolfe, G.W. 1979. A zoogeographic and taxonomic analysis of the Dytiscidae of Tennessee with an emphasis on the pulcher-undulatus species group of Hydroporus (Adephaga: Coleoptera). Ph.D. Dissertation. The University of Tennessee, Knoxville, TN. 153 pp.

Wolfe, G.W. 1980. The larva and pupa of Acilius fraternus fraternus (Coleoptera: Dytiscidae) from the Great Smoky Mountains, Tennessee. Coleopterists Bulletin 34: 121-126.

Woodroffe, G. E., and C. W. Coombs. 1961. A revision of the North American Cryptophagus Herbst (Coleoptera: Cryptophagidae). Miscellaneous Publications of the Entomological Society of America 2: 179-211.

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